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<lom:langstring xml:lang="x-none">10.3390/ani13091473</lom:langstring>

  
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<lom:langstring xml:lang="en">Equine Social Behaviour: Love, War and Tolerance</lom:langstring>

  
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<lom:langstring xml:lang="en">Sociality is an ethological need of horses that remained unchanged by domestication. Accordingly, it is essential to include horses&#39; social behavioural requirements and the opportunity to establish stable affiliative bonds in equine management systems and welfare assessment. Thus, this systematic review aims to provide an up-to-date analysis of equine intraspecific social ethograms. A literature review yielded 27 papers that met the inclusion criteria by studying adult (≥2 years) equine social behaviour with conspecifics using a well-defined ethogram. Social interactions were observed in 851 horses: 320 (semi-)feral free-ranging, 62 enclosed (semi-)feral and 469 domesticated, living in groups averaging 9.1 (mean +/- 6.8 s.d., range: 2-33) horses. The ethograms detailed in these 27 studies included a total of 40 (mean: 12.8/paper, range: 2-23) social behaviours, of which 60% (24/40) were agonistic, 30% (12/40) affiliative, 7.5% (3/40) investigative and 2.5% (1/40) neutral. The 27 publications included 67.7% agonistic and only 26% affiliative, 5.1% investigative and 1.2% neutral social behaviours in their methodology, thus focusing predominantly on socio-negative interactions. The strong emphasis on agonistic behaviours in equine ethology starkly contrasts with the rare occurrence of agonistic behaviours in stable horse groups and the well-established importance of affiliative interactions for equine welfare. The nuanced and complex equine social behaviour requires refinement of the ethogram with a greater focus on affiliative, ambivalent and indifferent interactions and the role of social tolerance in equine social networks to advance equine welfare assessment.</lom:langstring>

  
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<lom:language>eng</lom:language>

  
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<lom:langstring xml:lang="en">Horses Equus-Caballus; Feral Horses; Reproductive Success; Przewalski Horses; Domestic Horses; Young Horses; Sorraia Horses; Paddock Size; Older-Adults; Herd</lom:langstring>

  
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<lom:datetime>2023-11-14T09:44:26.460Z</lom:datetime>

  
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N:Torres Borda;Laura;
FN:Laura Torres Borda
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<lom:vcard>BEGIN:VCARD
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N:Auer;Ulrike;
FN:Ulrike Auer
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<lom:vcard>BEGIN:VCARD
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N:Jenner;Florien;
FN:Florien Jenner
X-ORCID:https://orcid.org/0000-0002-6977-1984
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